Ecosystem Dynamics and Nutrient Cycles

  • Keystone species exert strong control on a community by their ecological roles, or niches.
  • In contrast to dominant species, they are not usually abundant in a community.
  • Ecosystem engineers (or “foundation species”) cause physical changes in the environment that affect community structure.
  • Beaver dams can transform landscapes on a very large scale.
  • The first law of thermodynamics states that energy cannot be created or destroyed, only transferred or transformed.
  • Energy enters an ecosystem as solar radiation, is conserved, and is lost from organisms as heat.
  • The second law of thermodynamics states that every exchange of energy increases the entropy of the universe.
  • In an ecosystem, energy conversions are not completely efficient; some energy is always lost as heat.
  • The law of conservation of mass states that matter cannot be created or destroyed.
  • Chemical elements are continually recycled within ecosystems.
  • Autotrophs build molecules themselves using photosynthesis or chemosynthesis as an energy source.
  • Heterotrophs depend on the biosynthetic output of other organisms.
  • Energy and nutrients pass from primary producers (autotrophs) to primary consumers (herbivores) to secondary consumers (carnivores) to tertiary consumers (carnivores that feed on other carnivores) and so on…
  • Detritivores, or decomposers, are heterotrophs that derive their energy from detritus, nonliving organic matter.
  • Prokaryotes and fungi are the main decomposers.
  • Primary production makes up the bottom of the food chain.
  • Total primary production is known as the ecosystem’s gross primary production (GPP).
  • GPP is measured as the conversion of energy from light (or chemicals) to the chemical energy of organic molecules per unit time.
  • Net primary production (NPP) is GPP minus energy used by autotrophs for respiration.
  • NPP is the amount of new biomass added in a given time period.
  • Only NPP is available to consumers.
  • Tropical rain forests, estuaries, and coral reefs are among the most productive ecosystems per unit area.
  • Marine ecosystems are relatively unproductive per unit area but contribute much to global net primary production because of their size.
  • Secondary production of an ecosystem is the amount of chemical energy in food converted to new biomass during a given period of time.
  • When a caterpillar feeds on a leaf, only about one-sixth of the leaf’s energy is used for secondary production.
  • Trophic efficiency is the percentage of production transferred from one trophic level to the next.
  • It is usually about 10%, with a range of 5% to 20%.
    • About 90% of energy is not transferred.
    • Approximately 0.1% of chemical energy fixed by photosynthesis reaches a tertiary consumer.
  • The rate of decomposition is controlled by temperature, moisture, and nutrient availability.
  • Nutrient cycles are called biogeochemical cycles because they involve both biotic and abiotic components.
  • The oceans contain 97% of the biosphere’s water; 2% is in glaciers and polar ice caps, and 1% is in lakes, rivers, and groundwater.
  • Photosynthetic organisms convert CO2 to organic molecules that are consumed by heterotrophs.
  • Carbon reservoirs include fossil fuels, soils and sediments, solutes in oceans, plant and animal biomass, the atmosphere, and sedimentary rocks.
  • Nitrogen is a component of amino acids, proteins, and nucleic acids.
  • The main reservoir of nitrogen is the atmosphere (N2), though this nitrogen must be converted to NH4+ or NO3 for uptake by plants, via nitrogen fixation by bacteria.
  • Some bacteria can also use NO2.
  • Bioremediation is the use of organisms to detoxify ecosystems.